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中国(虫齿)科((虫齿)总目:(虫齿)亚目)昆虫的系统分类研究

Systematics of Psocidae(Psocodea:Psocomorpha) from China

【作者】 刘露希

【导师】 刘志琦;

【作者基本信息】 中国农业大学 , 农业昆虫与害虫防治, 2014, 博士

【摘要】 虫齿科Psocidae隶属于(?)总目Psocodea(?)亚目Psocomorpha(?)次目Psocetae,是(?)总目昆虫中种类最丰富的类群之一,目前世界已知81属1210种,分属于2亚科,广泛分布于古北界、东洋界、非洲界、澳洲界、新北界以及新热带界各大动物地理区系中。我国(?)科昆虫资源也是十分丰富,现已有记录37属357种。本论文对我国(?)科昆虫进行了系统分类研究。文中介绍了世界和我国(?)科的分类研究概况,并介绍了(?)科各虫态的形态特征;简述了(?)科的生物学和生态学特征。运用PAUP4.0软件对我国(?)科的属间关系进行系统发育分析,初步揭示了(?)科的单系性和我国(?)科各属间的亲缘关系;对联(?)属Symbiopsocus的种间系统发育关系进行了分析和讨论,验证了部分种分类地位变动的合理性。同时对(?)科在世界各大区系的分布情况做了初步分析,结果表明东洋界种类最为丰富,其次为新热带界、古北界和新北界,非洲界和澳洲界种类较少,各属分布不均,各有其不同的分布特点,有世界性分布、亚世界性分布、多区分布、双区分布和单区分布五种类型,其中带麻(?)属Trichadenotecnum是最大的属,其种类数量占整个(?)科种类数量的16.6%。另外各属在我国的分布以东洋界的种类最多,在此提供了相关的地理分布图表。根据已有的标本和文献,对我国(?)科进行了系统的分类研究并对现有种类进行分类记述,共收录了我国(?)科昆虫5属59种,并对其进行了详细的描述及绘图,其中包括7新种,提出了10同物异名种以及5新组合。新种:叉联(?)Symbiopsocus furcatus Liu et Liu,2011,阳明(?)Symbiopsocus yangminus Liu et Liu,2014,屏边曲(?)Sigmatoneura pinbiensis Liu et Liu,2011,暗色曲(?)Sigmatoneura aquilis Liu et Liu,2011,圆突拟新(?)Neopsocopsis convexa Liu et Liu,2012,多齿枝虫齿Clematocenea spinosus Liu et Liu,2013,五齿黑麻虫齿Atrichadenotecnum multuspinum Liu et Liu,2013;同物异名种:钳拟新蜡Neopsocopsis obconica (Li,2002)(=粗角拟新(?)Neopsocopsis hirticornis (Reuters,1893)),棒拟新(?)Neopsocopsis clavata (Li,2002)=粗角拟新虫齿Neopsocopsis hirticornis (Reuters,1893)),梯拟新虫齿Neopsocopsis tetraedrica (Li,2001)(=长距拟新(?)Neopsocopsis longiptera Vishnyakova,1986),裂瓣拟新虫齿Neopsocopsis schizopetala (Li,1997)(=深色拟新(?)Neopsocopsis profunda (Li,1995)),黄色拟新虫齿Neopsocopsis flavae (Li,1995)(=深色拟新虫齿Neopsocopsis profunda (Li,1995)),披拟新(?)Neopsocopsis lanceolata (Li,2002)(=淡黄拟新虫齿Neopsocopsis flavida (Li,1989)),松拟新(?)Neopsocopsis pini (Li,1990)(=淡黄拟新(?)1Neopsocopsis flavida (Li,1989)),二星丽斑(?)Psocomesites bimaculatum Li,1995(=三叉黑麻(?)Atrichadenotecnum trifurcatum (Li,1993)),凹歧痣(?)Clematostigma excavata Li,2002(=三叉黑麻(?)Atrichadenotecnum trifurcatum (Li,1993)),五齿丽斑虫齿Psocomesites edentalum Li,2005(=三叉黑麻虫齿Atrichadenotecnum trifurcatum (Li,1993))。对观察过的种类进行了重新描述和绘图,并编制了蜡科的分属检索表和部分属的分种检索表。整理完成了世界和中国(?)科属种名录。

【Abstract】 The family Psocidae belongs to Psocodea, Psocoptera, Psocetae, is one of the largest groups of Psocodea, rich in the diversity of species and ecology. So far, nearly1210species in81genera are recorded in the world, widely distributing in Palaearctic, Oriental, Afrotropic, Australasia, Nearctic and Neotropical Regions. About357species in37genera are known from China. The present dissertation deals with the systematics of the family Psocidae from China.The research history of Psocidae at home and abroad is briefly introduced. The general morphology of Psocidae is outlined. The information on the biology of Psocidae is provided. The phylogenetic relationships of Chinese genera of Psocidae are analyzed with PA UP4.0, the results show that Psocidae is a monophyletic group; and the relationships among species in the genus Symbiopsocus are also demonstrated. The distribution patterns of Psocidae are discussed both on world and China scale. It shows that the Oriental Region holds the most abundant Psocidae species; Neotropic Region, Palaearctic Region and Nearctic Region have a fewer species distributed; the species distributed in Afrotropic Region and Australasia Region are nearly equal in number. Each genus has its own distribution pattern. Among genera, the genus Trichadenotecnum is the biggest one which has more than200species. China belongs faunisticlly to two zoogeographical regions:the Palaearctic Region and the Oriental Region. The species from the Oriental Region compose the principal part of Chinese fauna. The relative charts of zoogeographical distribution are provided in the thesis.In the taxonomic section,59species in5genera from China are described, including seven new species, as well as10new synonyms and5new combinations. New species:Symbiopsocus furcatus Liu et Liu,2011, Symbiopsocus yangminus Liu et Liu,2014, Sigmatoneura pinbiensis Liu et Liu,2012, Sigmatoneura aquilis Liu et Liu,2012, Neopsocopsis convexa Liu et Liu,2012, Clematocenea spinosus Liu et Liu,2013, Atrichadenotecnum multuspinum Liu et Liu,2013; new synonyms:Neopsocopsis obconica (Li,2002)(=Neopsocopsis hitricornis (Reuters,1893)), Neopsocopsis clavata (Li,2002)(=Neopsocopsis hitricornis (Reuters,1893)), Neopsocopsis tetraedrica (Li,2002)(=Neopsocopsis longiptera Vishnyakova,1986), Neopsocopsis schizopetala (Li,1997)(=Neopsocopsis profunda (Li,1995)), Neopsocopsis flavae (Li,1995)(=Neopsocopsis profunda (Li,1995)), Neopsocopsis lanceolata (Li,2002)(=Neopsocopsis flavida (Li,1989)), Neopsocopsis pini (Li,1990)(=Neopsocopsis flavida (Li,1989)), Psocomesites bimaculatum Li,1995(=Atrichadenotecnum trifurcatum (Li,1993)), Clematostigma excavata Li,2002(=Atrichadenotecnum trifurcatum (Li,1993)), Psocomesites edentalum Li,2005(=Atrichadenotecnum trifurcatum (Li,1993)). A key to the Chinese genera of Psocidae, and keys to the Chinese species of the above genera are presented. The world and Chinese catalogues of subfamilies, genera, and species are provided.

【关键词】 (?)总目(?)亚目(?)次目虫齿科系统学中国
【Key words】 PsocodeaPsocomorphaPsocetaePsocidaesystematicsChina
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